Crompton, A. The cranial morphology of a new genus and species of ictidosauran. Proc. Zool. Soc. Lond. 130, 183–216 (1958).
Kermack, K. A. & Mussett, F. The jaw articulation of the Docodonta and the classification of Mesozoic mammals. Proc. R. Soc. Lond. B 149, 204–215 (1958).
Simpson, G. G. Diagnosis of the classes Reptilia and Mammalia. Evolution 14, 388–392 (1960).
Barghusen, H. R. & Hopson, J. A. Dentary–squamosal joint and the origin of mammals. Science 168, 573–575 (1970).
Rowe, T. Definition, diagnosis, and origin of Mammalia. J. Vertebr. Paleontol. 8, 241–264 (1988).
Luo, Z.-X., Kielan-Jaworowska, Z. & Cifelli, R. L. In quest for a phylogeny of Mesozoic mammals. Acta Palaeontol. Pol. 47, 1–78 (2002).
Rawson, J. R. et al. Brazilian fossils reveal homoplasy in the oldest mammalian jaw joint. Nature 634, 381–388 (2024).
He, X. & Cai, K. The tritylodont remains from Dashanpu, Zigong. J. Chengdu Coll. Geol. Suppl. 2, 33–45 (1984).
Crompton A, & Parkyn, D. On the lower jaw of Diarthrognathus and the origin of the mammalian lower jaw. Proc. Zool. Soc. Lond. 140, 697–749 (1963).
Reed, D., Iriarte‐Diaz, J. & Diekwisch, T. A three dimensional free body analysis describing variation in the musculoskeletal configuration of the cynodont lower jaw. Evol. Dev. 18, 41–53 (2016).
Lautenschlager, S., Gill, P. G., Luo, Z.-X., Fagan, M. J. & Rayfield, E. J. The role of miniaturization in the evolution of the mammalian jaw and middle ear. Nature 561, 533–537 (2018).
Crompton, A. W. The evolution of the mammalian jaw. Evolution 17, 431–439 (1963).
Kermack, K. A., Mussett, F. & Rigney, H. W. The lower jaw of Morganucodon. Zool. J. Linn. Soc. 53, 87–175 (1973).
Bramble, D. M. Origin of the mammalian feeding complex: models and mechanisms. Paleobiology 4, 271–301 (1978).
Sues, H.-D. Skull and dentition of two tritylodontid synapsids from the Lower Jurassic of western North America. Bull. Mus. Comp. Zool. 151, 217–268 (1986).
Crompton, A. W. & Hylander, W. in The Ecology and Biology of Mammal-like Reptiles (eds Nicholas, H. et al.) 263–282 (1986).
Grossnickle, D. M., Weaver, L. N., Jäger, K. R. & Schultz, J. A. The evolution of anteriorly directed molar occlusion in mammals. Zool. J. Linn. Soc. 194, 349–365 (2022).
West-Eberhard, M. Developmental Plasticity and Evolution (Oxford Univ. Press, 2003).
Gilbert, S. F., Bosch, T. C. & Ledón-Rettig, C. Eco-Evo-Devo: developmental symbiosis and developmental plasticity as evolutionary agents. Nat. Rev. Genet. 16, 611–622 (2015).
Müller, G. B. Why an extended evolutionary synthesis is necessary. Interface Focus 7, 20170015 (2017).
Crompton, A. W. in Studies in Vertebrate Evolution (eds Joysey, K. A. & Kemp, T. S.) 231–251 (Oliver & Boyd, 1972).
Kemp, T. S. The Origin and Evolution of Mammals (Oxford Univ. Press, 2005).
Kermack, K. A. The interrelations of early mammals. Zool. J. Linn. Soc. 47, 241–249 (1967).
Gow, C. The dentitions of the Tritheledontidae (Therapsida: Cynodontia). Proc. R. Soc. Lond. B 208, 461–481 (1980).
Allin, E. F. & Hopson, J. A. in The Evolutionary Biology of Hearing (eds Webster, D. B. et al.) 587–614 (Springer, 1992).
Patterson, B. & Olson, E. C. A triconodontid mammal from the Triassic of Yunnan. In International Colloquium On The Evolution Of Lower And Non-specialized Mammals 129–191 (Koninklijke Vlaamse Academiie voor Wetenschappen, 1961).
Kermack, K. A., Mussett, F. & Rigney, H. W. The skull of Morganucodon. Zool. J. Linn. Soc. 71, 1–158 (1981).
Mao, F. et al. Fossils document evolutionary changes of jaw joint to mammalian middle ear. Nature 628, 576–581 (2024).
Fourie, S. The jaw articulation of Tritylodontoideus maximus. S. Afr. J. Sci. 64, 255–265 (1968).
Crompton, A. & Sun, A.-L. Cranial structure and relationships of the Liassic mammal Sinoconodon. Zool. J. Linn. Soc. 85, 99–119 (1985).
Crompton, A. W. in Functional Morphology in Vertebrate Paleontology (ed. Thomason, T.) 55–75 (Cambridge Univ. Press, 1995).
Jasinoski, S. C. & Chinsamy, A. Mandibular histology and growth of the nonmammaliaform cynodont Tritylodon. J. Anat. 220, 564–579 (2012).
Allin, E. F. Evolution of the mammalian middle ear. J. Morphol. 147, 403–437 (1975).
Sidor, C. A. Evolutionary trends and the origin of the mammalian lower jaw. Paleobiology 29, 605–640 (2003).
Bertossa, R. C. Morphology and behaviour: functional links in development and evolution. Philos. Trans. R. Soc. B 366, 2056–2068 (2011).
Newman, S. A. & Müller, G. B. Epigenetic mechanisms of character origination. J. Exp. Zool. 288, 304–317 (2000).
Clark, J. M. & Hopson, J. A. Distinctive mammal-like reptile from Mexico and its bearing on the phylogeny of the Tritylodontidae. Nature 315, 398–400 (1985).
Lopatin, A. & Agadjanian, A. A tritylodont (Tritylodontidae, Synapsida) from the Mesozoic of Yakutia. Dokl. Biol. Sci. 419, 279–282 (2008).
Matsuoka, H., Kusuhashi, N. & Corfe, I. J. A new Early Cretaceous tritylodontid (Synapsida, Cynodontia, Mammaliamorpha) from the Kuwajima Formation (Tetori Group) of central Japan. J. Vertebr. Paleontol. 36, e1112289 (2016).
Averianov, A. O. et al. A tritylodontid synapsid from the Middle Jurassic of Siberia and the taxonomy of derived tritylodontids. J. Vertebr. Paleontol. 37, e1363767 (2017).
Velazco, P. M., Buczek, A. J. & Novacek, M. J. Two new tritylodontids (Synapsida, Cynodontia, Mammaliamorpha) from the Upper Jurassic, southwestern Mongolia. Am. Mus. Novit. 2017, 3874 (2017).
Panciroli, E., Walsh, S., Fraser, N. C., Brusatte, S. L. & Corfe, I. A reassessment of the postcanine dentition and systematics of the tritylodontid Stereognathus (Cynodontia, Tritylodontidae, Mammaliamorpha), from the Middle Jurassic of the United Kingdom. J. Vertebr. Paleontol. 37, e1351448 (2017).
Mao, F., Zhang, C., Liu, C. & Meng, J. Fossoriality and evolutionary development in two Cretaceous mammaliamorphs. Nature 592, 577–582 (2021).
Sun, A. L. Skull morphology of the tritylodont genus Bienotheroides of Sichuan. Sci. Sin. B 27, 970–984 (1984).
Liu, L. et al. New discovery of tritylodontids from the Middle Jurassic in Yunyang area, Chongqing and their paleogeographic significance. J. Palaeogeogr. 26, 1–15 (2024).
Crompton, A. W. & Jenkins, F. A. Molar occlusion in Late Triassic mammals. Biol. Rev. 43, 427–458 (1968).
Crompton, A. W. Postcanine occlusion in cynodonts and tritylodontids. Bull. Br. Mus. (Nat. Hist.) 21, 27–71 (1972).
Kalthoff, D. C. et al. Complementary approaches to tooth wear analysis in Tritylodontidae (Synapsida, Mammaliamorpha) reveal a generalist diet. PLoS One 14, e0220188 (2019).
Hopkins, S. S. The evolution of fossoriality and the adaptive role of horns in the Mylagaulidae (Mammalia: Rodentia). Proc. R. Soc. B 272, 1705–1713 (2005).
Liu, J., Soares, M. B. & Reichel, M. Massetognathus (Cynodontia, Traversodontidae) from the Santa Maria Formation of Brazil. Rev. Bras. Paleontol. 11, 27–36 (2008).
Hopson, J. A. & Kitching, J. W. A revised classification of cynodonts (Reptilia; Therapsida). Palaeontol. Afr. 14, 71–85 (1972).
Hopson, J. & Crompton, A. in Evolutionary Biology (eds Dobzhansky, T. et al.) 16–72 (Appleton-Century-Crofts, 1969).
Anthwal, N. & Tucker, A. S. Evolution and development of the mammalian jaw joint: Making a novel structure. Evol. Dev. 25, 3–14 (2023).
Hunter, J. P. & Jernvall, J. The hypocone as a key innovation in mammalian evolution. Proc. Natl Acad. Sci. USA 92, 10718–10722 (1995).
Donoghue, M. J. & Sanderson, M. J. Confluence, synnovation, and depauperons in plant diversification. New Phytol. 207, 260–274 (2015).
Erwin, D. H. A conceptual framework of evolutionary novelty and innovation. Biol. Rev. 96, 1–15 (2021).
Müller, G. B. & Wagner, G. P. Novelty in evolution: restructuring the concept. Annu. Rev. Ecol. Syst. 22, 229–256 (1991).
Arthur, W. Intraspecific variation in developmental characters: the origin of evolutionary novelties. Am. Zool. 40, 811–818 (2000).
Martin, T. & Rauhut, O. W. M. Mandible and dentition of Asfaltomylos patagonicus (Australosphenida, Mammalia) and the evolution of tribosphenic teeth. J. Vertebr. Paleontol. 25, 414–425 (2005).
Morales-García, N. M., Gill, P. G., Janis, C. M. & Rayfield, E. J. Jaw shape and mechanical advantage are indicative of diet in Mesozoic mammals. Commun. Biol. 4, 242 (2021).
Debuysschere, M., Gheerbrant, E. & Allain, R. Earliest known European mammals: a review of the morganucodonta from Saint-Nicolas-de-Port (Upper Triassic, France). J. Syst. Palaeontol. 13, 825–855 (2015).
Gill, P. G. et al. Dietary specializations and diversity in feeding ecology of the earliest stem mammals. Nature 512, 303–305 (2014).
Sues, H.-D. The relationships of the Tritylodontidae (Synapsida). Zool. J. Linn. Soc. 85, 205–217 (1985).
Cooper, K. L. The case against simplistic genetic explanations of evolution. Development 151, dev203077 (2024).
van Valen, L. Festschrift: Evolutionary Biology. Vol. 6. Theodosius Dobzhansky, Max K. Hecht and William C. Steere, eds. Science 180, 488 (1973).
Simpson, G. G. A Catalogue of the Mesozoic Mammalia in the Geological Department of the British Museum (Trustees of the British Museum, 1928).
Setoguchi, T., Matsuda, M. & Matsuoka, H. New discovery of an Early Cretaceous tritylodontid (Reptilia, Therapsida) from Japan and the phylogenetic reconstruction of Tritylodontidae based on the dental characters. In Seventh Annual Meeting of the Chinese Society of Vertebrate Paleontology (eds Wang Y. Q. & Deng, T.) 117-124 (China Ocean Press, 1999).
Matsuoka, H. in Fossils of the Kuwajima “Kaseki-kabe” (Fossil-bluff): Scientific Report on a Neocomian (Early Cretaceous) Fossil Assemblage from the Kuwajima Formation, Tetori Group, Shiramine, Ishikawa, Japan (ed. Matsuoka, H.) 53–74 (Shiramine Village Board of Education, 2000).
Watabe, M., Tsubamoto, T. & Tsogtbaatar, K. A new tritylodontid synapsid from Mongolia. Acta Palaeontol. Pol. 52, 263–274 (2007).
Davis, B. M., Jäger, K. R. K., Rougier, G. W., Trujillo, K. & Chamberlain, K. A morganucodontan mammaliaform from the Upper Jurassic Morrison Formation, Utah, USA. Acta Palaeontol. Pol. 67, 77–93 (2022).
Mao, F. et al. Jurassic shuotheriids show earliest dental diversification of mammaliaforms. Nature 628, 569–575 (2024).
Mao, F. et al. Convergent evolution of diverse jaw-joints in mammaliamorph. Zenodo https://doi.org/10.5281/zenodo.16193266 (2025).
Luo, Z.-X. Developmental patterns in Mesozoic evolution of mammal ears. Annu. Rev. Ecol. Syst. 42, 355–380 (2011).
Soares, M. B., Schultz, C. L. & Horn, B. L. New information on Riograndia guaibensis Bonaparte, Ferigolo & Ribeiro, 2001 (Eucynodontia, Tritheledontidae) from the Late Triassic of southern Brazil: anatomical and biostratigraphic implications. An. Acad. Bras. Cienc. 83, 329–354 (2011).
Rodrigues, P. G. et al. Digital cranial endocast of Riograndia guaibensis (Late Triassic, Brazil) sheds light on the evolution of the brain in non-mammalian cynodonts. Hist. Biol. 31, 1195–1212 (2019).
Kerber, L. et al. An additional brain endocast of the ictidosaur Riograndia guaibensis (Eucynodontia: Probainognathia): intraspecific variation of endocranial traits. An. Acad. Bras. Cienc. 93, e20200084 (2021).
Mao, F. & Meng, J. A new haramiyidan mammal from the Jurassic Yanliao Biota and comparisons with other haramiyidans. Zool. J. Linn. Soc. 186, 529–552 (2019).
Meng, J., Wang, Y. & Li, C. Transitional mammalian middle ear from a new Cretaceous Jehol eutriconodont. Nature 472, 181–185 (2011).
Mao, F. et al. Integrated hearing and chewing modules decoupled in a Cretaceous stem therian mammal. Science 367, 305–308 (2020).
Mao, F., Liu, C., Chase, M. H., Smith, A. K. & Meng, J. Exploring ancestral phenotypes and evolutionary development of the mammalian middle ear based on Early Cretaceous Jehol mammals. Natl Sci. Rev. 8, nwaa188 (2021).
